XI B RARY OF THE UNIVERSITY or ILLINOIS 590.5 FX V. 37-38 BtOLOQi ^test Date stamped below AR04 15 L161 O-1096 Bats from Guerrero^ Jalisco and Oaxaca^ Mexico Luis de la Torre Associate, Division of Mammals This paper is based on a small collection of bats made by the author in Guerrero and Oaxaca from July 22 to August 5, 1952, and in Jalisco from December 7 to 11, 1954. The primary objective of this collecting was to obtain additional specimens of Sturnira to be used for a systematic review of that genus. In sampling caves and other likely habitats, representatives of fourteen species of bats were collected, including the first series to be taken of Enchisthenes harti. I am indebted to Miss Barbara Lawrence, Museum of Com- parative Zoology, and to Dr. E. Raymond Hall, Museum of Natural History, University of Kansas, for the loan of specimens in their care; to Mr. R. W. Hayman, British Museum (Natural History), for comparing specimens of Chilonyderis with the types of C. psilotis; and to Mr. and Mrs. Loren P. Woods, Chicago Natural History Museum, for their kindness in making possible my visit to the interesting Jalisco region. The patient and enthusiastic help given by my wife, June, in many capacities is also gratefully acknow- ledged. LIST OF SPECIES With the exceptions noted, the specimens from Guerrero and Oaxaca were taken in caves. Those from Ciudad Guzmdn, Jalisco, were netted at night in a grove of native, small-fruit Ficus trees. The measurements given are in millimeters and, unless other- wise indicated, apply to the entire series. The average measurement, when given, is followed by the extremes in parentheses. Balantiopteryx plicata plicata Peters Balantiopteryx plicata Peters, 1867, Monatsb, Preuss. Akad. Wiss. Berlin, 1867: 476. 696 696 FIELDIANA: ZOOLOGY, VOLUME 37 Type ZocaZi^^/.^Puntarenas, Puntarenas, Costa Rica. Specimens collected. Total 32 (2 skins with skulls, 30 in alcohol) . Guerrero: 5 mi. NW. of Acapulco (12 males, 14 females); 12 mi. S. of Mexcala (1 female) . Oaxaca : Tehuantepec (5 young females) . Measurements. Forearm of males from Acapulco 41.3 (40.0- 43.0), females 42.6 (40.0-46.0); female from Mexcala 41.8. Chilonycteris rubiginosa mexicana Miller Chilonyderis mexicana Miller, 1902, Proc. Acad. Nat. Sci. Philadelphia, 54: 401. C[hilonycteris] p[arnellii] mexicana, Koopman, 1955, Jour. Mamm., 36: 112, Type locality. San Bias, Nayarit, Mexico. Specimens collected. Total 8 (2 skins with skulls, 6 in alcohol). Guerrero: 5 mi. NW. of Acapulco (7 males, 1 female). Measurements. Forearm of males 57.3 (55.7-58.8), female 59.5. Remarks. It cannot be established that parnelli Gray 1843 has priority over rubiginosa Wagner 1843. The issue of the Proceedings of the Zoological Society of London where the name parnelli first appears is dated April 11, 1843, as shown by Koopman (Jour. Mamm., 1955, 36: 111), but the actual date of publication is October of that year (Duncan, Proc. Zool. Soc. London, 1937: 79). The name rubiginosa appears in the Archiv fiir Naturgeschichte after a paper read in August, but the exact date of publication is not known. In the absence of conclusive evidence, the better known and more widely used name rubiginosa should be retained. Chilonycteris psilotis Dobson Chilonycteris psilotis Dobson, 1878, Cat. Chirop. Brit. Mus., p. 451. Miller, 1902, Proc. Biol. Soc. Washington, 15: 249 Isthmus of Tehuantepec, Mexico. Chilonycteris personata, J. A. Allen (not Wagner), 1911, Bull. Amer. Mus. Nat. Hist., 30: 263. G. M. Allen, 1935, Jour. Mamm., 16: 227. Dalquest and Hall, 1949, Jour. Mamm., 30: 425. Davis and Russell, 1952, Jour. Mamm., 33: 234; 1954, op. cit., 35: 67. Chilonycteris torrei continentis Sanborn, 1938, Occ. Pap. Mus. Zool., Univ. Michigan, no. 373: 1 Laguna de Zotz, Peten, Guatemala (type locality). Type locality. Here designated as Tehuantepec, Oaxaca, Mexico. Specimens collected. Total 3 (2 skins with skulls, 1 in alcohol). Guerrero: 5 mi. NW. of Acapulco (3 males). Measurements. Forearm 42.5, 43.2, 44.0. DE LA TORRE: BATS FROM MEXICO 697 Remarks. These specimens agree with the type of C. torrei con- tinentis Sanborn, and with Guatemala specimens that are identical with the types of psilotis. Additional representatives of psilotis have been examined from Mexico, Central America, and Venezuela. Inasmuch as it has not been demonstrated that C. psilotis and the Brazilian C. personata are conspecific, the name psilotis, rather than personata, should be used for the small, mainland Chilonycteris. Pteronotus davyi fulvus Thomas Chilonycteris davyi fulvus Thomas, 1892, Ann. Mag. Nat. Hist., (6), 10: 410. Type locality. Las Peiias, Jalisco, Mexico. Specimens collected. Total 11 (in alcohol). Guerrero: 5 mi. NW. of Acapulco (9 males, 2 females). Measurements. Forearm, males 44.1 (43.2-45.5), females 44.0, 45.0. Mormoops megalophylla megalophylla Peters Mormops megalophylla Peters. 1864, Monatsb. Preuss. Akad. Wiss. Berlin, 1864:381. Type locality. Southern Mexico. Specimens collected. Total 27 (in alcohol). Guerrero: 5 mi. NW. of Acapulco (27 males, no females found). Measurements. Forearm 54.9 (53.0-59.0). Micronycteris megalotis mexicana Miller Micronycteris megalotis mexicana Miller, 1898, Proc. Acad. Nat. Sci. Phila- delphia, 50: 329. Type locality. 15 mi. S. of Ciudad Guzman, Platanar, Jalisco, Mexico. Specimens collected. 1 (in alcohol). Jalisco: 2 mi. N. of Ciudad Guzmdn (1 female). Measurements. Forearm 36.3. Glossophaga soricina leachi Gray Monophyllus leachii Gray, 1844, Voyage of the Sulphur, Zool., 1: 18. Type locality. Realejo, Nicaragua. Specimens collected. Total 9 (in alcohol). Guerrero: 5 mi. NW. of Acapulco (6 males, 3 females). Measurements. Forearm 36.3 (35.0-37.5). 698 FIELDIANA: ZOOLOGY, VOLUME 37 Remarks. One specimen was taken in a mouse trap baited with ripe banana, Leptonycteris nivalis nivalis Saussure M[=Ischnoglossa] nivalis Saussure, 1860, Rev. Mag. Zool., Paris (2), 12: 492. Leptonycteris nivalis yerhabuenae Martinez and Villa, 1940, An. Inst. Biol. Univ. Nac. Mexico, 11: 313 Yerbabuena, Guerrero, Mexico (type locality). Type locality. Near snow line on Mount Orizaba, Veracruz, Mexico. Specimens collected. 1 (in alcohol). Guerrero: 12 mi. S. of Mexcala (1 male). Measurements. Forearm 54.0. Remarks. This specimen, a topotype of yerbabuenae, is in- distinguishable from a topotype of true nivalis from Orizaba. In size, it also agrees with specimens from Colima, but is smaller than those from Nuevo Leon, Texas, and Arizona. A size cline from North to South is indicated. Artibeus jamaicensis jamaicensis Leach Ariibeus jamaicensis Leach, 1821, Trans. Linn. Soc. London, 13: 75. Type locality. Jamaica. Specimens collected. Total 5 (in alcohol). Jalisco: 2 mi. N. of Ciudad Guzman (3 males, 2 females). Measurements. Forearm 58.0 (55.5-60.0). Remarks. All specimens in the series have the third upper molar. Artibeus cinereus toltecus Saussure Stenoderma tolteca Saussure, 1860, Rev. Mag. Zool., Paris, (2), 12: 427. Type locality. Mirador, Veracruz, Mexico. Specimens collected. Total 8 (in alcohol). JALISCO: 2 mi. N. of Ciudad Guzman (5 males, 3 females). Measurements. Forearm 40.0 (37.0-42.0). Remarks. The palate in two individuals appeared abnormally wide, and the arrangement of the upper teeth (pi. 30, fig. 1) differed significantly from the normal condition in Artibeus cinereus (pi. 30, figs. 4, 5, 6) . Study of the skulls revealed that they were of extreme age, and the labial edge of the maxillary bone had been eroded (pi. 30, fig. 2) leaving a good part of the roots unprotected. It is probable that if this weakened condition develops early in life, the Fig. 146. a, View from the Ciudad Guzmdn basin to the southwest. Volcan de Colima in the distance, b, View of basin showing scattered fig trees frequented by fruit-eating bats, c, Eastern edge of basin showing feeding habitat of Enchis- thenes harti, Artibeus j. jamaicensis, A. cinereus toltecus, Sturnira lilium parvidens, and Micronycteris megalotis mexicana. 699 700 FIELD lANA: ZOOLOGY, VOLUME 37 lateral mastication movements very likely would alter the align- ment of the teeth and cause them to flare out as in this case (pi. 30, fig. 3). Enchisthenes harti Thomas. Plate 31. Artibeus harti Thomas, 1892, Ann. Mag. Nat. Hist., (6), 10: 408. Enchisthenes harti, Sanborn, 1932, Ann. Carnegie Mus., 21: 179 Venezuela (skin and skull); Goodwin, 1940, Amer. Mus. Nov., no. 1075: 2 Hon- duras (skin and skull); Ecuador (incomplete skulls from owl pellets); Goodwin, 1954, Amer. Mus. Nov., no. 1689: 5 Tamaulipas, Mexico (incomplete skull). Type locality. Trinidad, British West Indies. Specimens collected. Total 12 (1 skin with skull, 11 in alcohol). Jalisco: 2 mi. N. of Ciudad Guzman (8 males, 4 females). Measurements. Forearm 39.8 (38.2-43.1); greatest length of skull, of a male and two females 20.4, 20.9, 20.9; condylobasal length 18.8, 19.0, 19.0; zygomatic breadth 12.1, 12.2, 12.1; mastoid breadth 10.8, 10.7, 10.6; palatal length 9.0, 9.8, 9.1; postorbital constriction 5.4, 5.6, 5.3; maxillary tooth row 6.8, 7.0, 6.9; greatest width across molars 8.4, 8.6, 8.3; length of mandible 13.1, 13.4, 13.9; lower canine to condyle 12.9, 13.2, 13.5; mandibular tooth row 7.3, 7.5, 7.3. Remarks. E. harti, previously known from only three complete specimens, was common in the Ficus groves (fig. 146, c) near Ciudad Guzman. This village lies in a basin rimmed by the Sierra Nevada de Colima on the west (fig. 146, b), and by lower, pine covered hills on the east (fig. 146, c). The feeding habits of Enchisthenes, judging from the individuals observed, probably make it a rather easy prey of owls. The fruit eaten in this area is small, about a centimeter in diameter. It is quickly snipped from the tree in flight, and carried to a convenient branch where it is eaten. The forearms are used to handle and turn the fruit as parts of it are consumed, and it is in these movements that the exposed bat probably is seen by its predators. It is interesting to note that E. harti and Sturnira lilium parvidens, both tropical species, were taken on one occasion when the tem- perature was only 36 F. Desmodus rotundus murinus Wagner D[esmodus] murinus Wagner, 1840, Schreber's Saugth., Suppl., 1: 377. Type locality. Mexico. DE LA TORRE: BATS FROM MEXICO 701 Specimens collected. Total 2 (in alcohol). Guerrero: 5 mi. NW. of Acapulco (1 male, 1 female) . Measurements. Forearm, male 58.2, female 60.6. Molossus sinaloae Allen Molossiis sinaloae Allen, 1906, Bull. Amer. Mus. Nat. Hist., 22: 236. Type locality. Esquinapa, Sinaloa, Mexico. Specimens collected. Total 9 females (in alcohol). Guerrero: 15 mi. NW. of Acapulco. Measurements. Forearm 47.9 (46.5-49.8). Remarks. These specimens were taken under the dry fronds of a coconut palm. Miller and Kellogg (1955, U. S. Nat. Mus. Bull. 205: 119) follow G. M. Allen in regarding sinaloae a race of rufus Geoffroy. M. sinaloae, however, is a medium size species and is distinct from the larger M. rufus of which nigricans Miller is the northern race. M. r. nigricans and M. r. rufus differ only in average size, while M. sinaloae can always be separated from M. rufus on absolute characters. Fieldiana: Zoology, Volume 37 Plate 30 Palate, maxilla and rostrum of Artibeus cinereus toUecus: 1-3, distorted condition in female of extreme age; 4-6, normal condition in adult female. X 4. Fieldiana: Zoology, Volume 37 Plate 31 Enchisthenes harti: 1, mandible; 2, ventral surface of skull; 3, dorsal surface of skull. X 4. Index Principal discussion and all new names indicated by bold-faced type. Abo Formation 389 Abraeinae 605, 610 Abraeus 601 acicularis 620 phyllobius 601 punctiger 624 vividulus 601 abruptum, Platysoma 629 aburri, Aburria 21 Aburria aburri 21 acanthognathus, Lophodolus 299 Acanthotermes 468, 486, 512 Achalinus spinalis 174 acicularis, Abraeus 620 Bacanius 605, 606, 620 Aciculitermes 515 acme, Bulimulus 325 Acorhinotermes 469, 493, 509 Amfus 602, 605 biloculatus 612 caledoniae 605, 607, 612, 614 lilliputianus 612 schmidti 605, 607, 612, 614 actinopterygians 388 Actitis macularia 154 aculeatus, Argyropelecus 275 acuminata, Erolia 154 acunhae, Nesospiza 147, 152 horny palate 151 acutirostris, Calamaria 199 Adioryx 126 Holocentrus 92, 109 Admiral Formation 389 adusta, Nenia 305 tutnens, Nenia 305 aegyptiaca, Tadarida 595 aegyptiacus, Rousettus 595 Aeie/es 620 crenatus 605, 606, 610 daubani 610 iaetns 610 longipes 614 politus 610 aequatorialis, Penelope 17 aequinoctialis, Succinea 305 Afrosubulitermes 515 agilis, Mus 648 Agouti paca 215 Ahamitermes 469, 503, 510 Ailuropoda 44 Aistopoda 531 AA;odon aerosMS 234 berlepschii 231 jelskii 254 pulcherrimus 227 pyrrhotis 228 nigrita 237 akodoni, Amblyopinus 228 alatus, Lampanyctus 289 ai6a, Egretta 153 Monopterus 53, 59 Muraena 59 alberti, Crax 17 albinism, in cracids 19 albiventer, Calamaria 199 albopunctata, Calamaria 199 alidae, Calamaria 200 Allodontermes 473, 513 Allognathotermes 511 alstoni, Neotomodon 547 alticolus, Edrabius 259 amabilis, Argyropelecus 276 amazonicus, Holochilus 659, 667 brasiliensis 667, 669 Ambassis, ecological distribution 81 gymnocephalus 52, 54, 71 interrupta 52flf, 71 ecological distribution 82 food 71, 84 nalua 53, 72 predator of 68 Amblyopininae 21 Iff Amblyopinini 221 Amblyopinodes 216, 218, 220, 222, 247 claviger 249 gahani 248 peruvianas 250 piceus 248 sitnilis 251 tvdvdssos'i 249 Amblyopinus 212, 214, 216-18, 220, 221 akodoni 228 ancashi 232 angustus 247 bequaerti 247 brandesi 227 chilomysi 239 colombiae 231 emarginatus 239 fuegensis 216, 237 705 706 FIELDIANA: ZOOLOGY, VOLUME 37 henseli 212, 247 hershkovitzi 238 huilae 242 jelskii 212, 224 kalinowskii 243 kofordi 231 longus 234 marmosae 247 tnonticolus 228 montivagans 240 oryzomysi 240 pacae 244 punae 227 sanborni 238 schmidti 231 spinipennis 234 waterhousei 219, 243 amblyurus, Hemirhamphus 67 Zenarchopterus 52ff, 67 food 67, 84 americanus, Ursus 25 Amitermes 467, 469, 472, 473, 477, 480, 483, 485, 488, 491, 493, 501, 503, 510 Amitermitinae 493, 505, 506, 510, 516, 562 amoenus, Neomyripristis 94 Amphibia, apsidospondylous 388 lepospondylous 388 Amphicynodontinae 44 amplus, Peromyscus difficilis 5^7 Ampullariidae 303 Ampullarius {Ampullarius) bridgesii 304 insularum 304 papyraceus 303 (Limnopomus) crassus 303 nubilus 303 Anabantoidea, ecological distribution 83 Anacanthotermes 473, 482, 509 anale, Scopelarchus 284 Anapleus 601 Anatidae 153 ancashi, Amblyopinus 232 Ancistrotermes 513 andium, Radiodiscus 308 andivaga, Streptaxis 329 Systrophia 308 andoicus, Bulimulus 324 Angelosaurus 395, 398 angouya, Mus 660 Oryzomys 654 angrandi, Bulimulus 315 kalinowskii, Nenia 306 Nenia 306 urubambensis, Nenia 306 Anyui^a borneensis 52ff, 58 ecological distribution 82 food 58, 84 predator of 62 Angularitermes 469, 495, 497, 514 Angulitermes 471, 485, 501, 512 angustus, Amblyopinus 247 anguya, Holochilus 647 Mus ( Holochilus) 648, 660, 666 Anodontostoma chacunda 52, 53, 55 Anomodontia 387, 398 distribution, geographic 390 geologic 390 Anoplogaster cornuta 292 Anoplogastridae 292 Anoplotermes 472, 484, 491, 493, 500, 510 Anthias maculatus 73 apaporiensis, Caiman sclerops 340 Apicotermes 473, 485, 511 Apilitermes 511 Apobletes 628 appendiculata 627 montrouzieri 605, 607, 627 Apocryptes brachypterus 78 riduosus 80 Apoda 531 Apodes, ecological distribution 83 Aporodon 644 appendiculata, Apobletes 627 apraeocularis, Calamaria 200 arborifera, Linophryne 300 Archotermopsis 468, 470, 482, 507, 509 Arctibeus lineatus 409 Arctotherium 25 arenarius, Miniopterus natalensis 595 Argfas 579ff 6oMeii 580, 594, 598 (Chiropterargas) boueti 580 confusus 579, 586, 594, 598 vespertilionis 586, 596, 598 Argasidae 579 argenteus, Diretmus 292 argentimaculata, Sciaena 73 argentimaculatus, Lutianus 54, 55, 73 food 84, 85 argentinus, Edrabius 258 argus, Chaetodon 75 Scatophagus 52, 54, 55, 75 argyromus, Myripristis 136 Argyropelecus aculeatus 275 amabilis 276 hemigymnus 274 Ariidae, ecological distribution 86 Ariophantidae 308 Arms caelatus 64 fenosMs 65 arkansas, Catapleura 377 armatus, Neoniphon 94 Armitermes 495, 514 Arroyo Formation 389 fauna 390flf artensis, Saprinus 605, 608, 609 arthuri-muelleri, Rugitermes 576 Ariibetts cinereus toltecus 698 /larii 700 jamaicensis jamaicensis 698 vittatus 405 INDEX 707 ascendens, BuUmulus derelidus 317 ascensionis, Holocentrus 93, 100, 106, 122, 134 Perca 100 Asellia tridens tridens 595 assimilis, Carcinops 619 Astronesthes leucopogon 278 Astronesthidae 278 atahualpa, Epiphragmophora 327 atavus, TreUmys 646 ataxus, BuUmulus 320 ater, Lampanyctus 288 atlantica, Evermannella atrata 284 atlanticus, Diogenichthys 286 Tetragonurus 298 Atianfma 158, 159 rogersi 142 Atractaspis 168 Afracitts 168 atradus, Beloholocentrus 95 atrata atlantica, Evermannella 284 atrum, Cyema 291 attenuata, Vinciguerria 273 auritus christiei, Plecotus 595 Reithrodon 646 australasiae, Saprinus 609 Australian region, termite fauna of 470, 476, 508flf Attsirah'terTwes 469, 477, 503, 515 Autunian fauna 389ff Bacanius 602, 605 acicularis 605, 606, 620 fauveli 605, 622, 624, 625 hamatus 620 pundiger 605, 606, 622, 624, 625 remingtoni 605, 606, 622, 624 sulcisternus 620 Bagrus nemurus 65 Balantiopteryx plicata plicata 695 6aZ6o, Evermannella 284 balnearum, Holochilus 656, 661, 665 Holochilus brasiliensis 665 Barbodes sealei 62 Barbus elongatus 62 Bartramia longicauda 154 Basidentitermes 512 Bathophilus metallicus 281 Bathylagidae 284 Bathylagus glacialis 284 bathyphila, Lampadena 288 bathyphilum, Gonostoma 272 bats, ticks of 579ff Beanea 95 trivittata 95 6eani, Serrivomer 291 beccari, Calamaria 200 Belle Plains Formation 389 Beloholocentrus 95 atradus 95 Belone strongylura 66 benjaminsi, Calamaria 200 benoiti, Hygophum 285 Benthodesmus simonyi 298 bequaerti, Amblyopinus 247 berbicensis, Holochilus 661 Holochilus brasiliensis 655, 669 berlepschi, Crypturellus cinereus 15 Beryciformes 121 Beto unimxiculata 53, 54, 70 ecological distribution 82, 86 food 70, 84 bibulbosus, Eustomias 282 bicolor, Calamaria 192, 200 bicornis, Ceratostethus 52, 54fif, 68 Linophryne 300 Neostethus 68 bidentatus, Urotriainus 563, 564 bigelowi, Eustomias 282 bikolana, Vaimosa 79 bikolanus, Pseudogobius 52, 54, 79 biloculatus, Acritus 612 bitaeniatus, Thaumastus 309 bitorques, Calamaria 200 blakfordi, Yarella 272 boa ferox, Stomias 278 boddaerti, Boleophthalmus 55, 80 food 80, 85 GobiMs 80 Boleophthalmus boddaerti 55, 80 food 80, 85 bonapartei, Nothocercus 12 Bonapartia pedialota 269 borneensis, Anguilla 52flf, 58 ecological distribution 82 food 58, 84 Calamaria 177ff, 194, 200 Gastromyzon 64 Neostethus 52, 54ff, 69 Borneo, North, fishes of 47ff Borneo Zoological Expedition 47flF boucardi, Crypturellus 16 boueti, Argas 580, 594 6o2/Ki levipes, Peromyscus 545 brachygnathus, Rhinolophus clivosus 595 Brachygobius doriae 52, 77 food 77, 85 brachypterus, Apocryptes 78 Holocentrus 109 Gobiopterus 55, 78 brachyrhynchus, Rhynchichthys 93 brachyura, Calamaria 192, 200 brachyurus, Doryichthys 55, 67 Syngnathus 67 brandesi, Amblyopinus 227 brasiliensis amazonicus, Holochilus 667 balnearum, Holochilus 665 berbicensis, Holochilus 655, 669 brasiliensis, Holochilus 662 guianae, Holochilus 668 Hesperomys (Holochilus) 666 Holochilomy 662 Holochilus 645, 648, 653, 654, 657, 658, 660, 661, 666 708 FIELDIANA: ZOOLOGY, VOLUME 37 incarum, Holochilus 668 leucogaster, Holochilus 648, 665 Mus 660, 662, 663, 667 nanus, Holochilus 666, 669 vulpinus, Holochilus 661 bratieri, Cyclothone 272 Brazil, shells from 303flf brevibarbatus, Stomias 279 brevidentatus, Serrivomer 291 bridgesii, Ampullarius 304 bridwelli, Spalacotheroides 690 brooki, Calamaria 179 bruegeli, Calamaria 185, 187 brunnescens, Penelope 17 Bubulcus ibis 153 Imccinatus, Oryzomys 654, 660 Buchen, Walter 140 buchi, Calamaria 195, 200 buckleyi, Odontophorus gujanensis 21 Bulbitermes 481, 514 Buliminus aegyptiacus 429, 432 chondriformis 419 heptodon 429 septemdentalus 422 sexdentatus 436 Bulimulidae 309, 331 Bulimulus acme 325 andoicus 324 angrandi 315 (Atax5) infundibulum 320 infumlibulum umbilicatellus 320 perforatum 320 cereicola 317 culmineus 315 dentritis 319 derelictus 317 eJafus 324 extensus 323 (Geoceras) veruculum 320 (Geopyrgus) turritus 336 hamiltoni 325 infundibulum 320 umbilicatellus 320 jocosensis 322 (Lissoacme) cereicola 317 dentritis 319 derelictus ascendens 317 Tnodestus 318 reconditus 320 rudistriatus 318 scutulatus 319 longitudinalis 316 modestus 318 mutabilis 334 n^TTiorensis 316 obeliscus 322 (Peronaetis) acme 325 andotcus 324 eiau